DURHAM, N.C. — Riley Leonard ran for one touchdown and threw for another as the Duke Blue Devils upset No. 9 Clemson, 28-7, on Monday night, capping a wild Week 1 of college football.

The Tigers (0-1) outgained the Blue Devils (1-0) and won the time of possession battle, but three turnovers doomed the new-look Clemson offense.

Cade Klubnik, entering his first full season as Clemson’s started quarterback, was intercepted, and the Tigers also lost two fumbles in the loss.

Jaquez Moore and Jordan Waters authored touchdown runs in support of Leonard, and the Blue Devils, who finished 9-4 last season including a bowl win over UCF, and have now won three straight, dating back to Nov. 26.

Clemson, last season’s ACC champions who lost to Tennessee in the Orange Bowl to end the year, revamped their offense in the offseason, bringing in Garrett Riley as the new coordinator.

Last season, Riley orchestrated the TCU offense, helping the Horned Frogs to the College Football Playoff title game. This was not the beginning coach Dabo Swinney drew up, when he hired Riley.

The Associated Press contributed to this report.